MESOCERATOCANTHUS FOSSILS FROM THE EARLY CRETACEOUS OF CHINA REINTERPRETED AS THE EARLIEST PASSALOIDS (COLEOPTERA: SCARABAEOIDEA)
DOI:
https://doi.org/10.5710/AMGH.26.04.2022.3502Keywords:
Scarabaeoidea, Hybosoridae, Passalopalpidae, Passalidae, China, Yixian FormationAbstract
The scarabaeoid beetle Mesoceratocanthus from the Early Cretaceous Yixian Formation was originally placed in the extant tribe Ivieolini (Hybosoridae). Our re-examination of the holotype and a new specimen demonstrates that Mesoceratocanthus does not possess the diagnostic characters of Ivieolini or Hybosoridae. Instead, it should be assigned to the passaloid lineage (Passalopalpidae + Passalidae) and appears closely related to the extinct family Passalopalpidae. The revised placement of Mesoceratocanthus further extends the earliest occurrence of the passaloid clade (~125 Ma, Barremian to early Aptian).

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